教育背景

1990.09-1995.07 清华大学 工程物理系 本科 工学学士

2000.09-2006.07 北京大学 环境学院 理学博士 

  工作履历

1995.07-1999.09 浙江秦山核电联营有限公司 生产部 助理工程师

1999.09-2000.07 山东青岛能电力设备有限公司 研发处 工程师

2006.07-2008.12 中国农业大学 资源与环境学院 助教/讲师

2008.12-2014.12 中国农业大学 资源与环境学院 副教授

2014.12-             中国农业大学 资源与环境学院 教授

  学术兼职

Journal of Plant Ecology编委（2020-）

《植物生态学报》编委（2014-）

国际生物地理学会会员（2018-）

中国地理学会生物地理学专业委员（2017-）

北京生态学学会监事（2017-）

国家现代海洋牧场科技创新联盟专家委员会（2017-）委员

欧洲EGU会员（2016-）

  研究领域

化学计量生态和生物化学地理；植物功能属性和全球环境变化

实验室目前重点研究我国北方和东部主要生态系统类型的化学计量特征及其与其他功能属性的内在联系，并探索在海陆、区域和局域尺度上这些特征和属性对气候（如变暖、干旱）和其他人类活动导致的全球性变化（如面源污染、地表类型改变、生物多样性损失等）的潜在响应。进一步可分为如下两个研究方向：

生态化学计量 研究C/N/P/S等生命基本元素在生态系统中的耦合关系，及其在生物圈特别是植被及环境（土-水-气）中的循环、转化与分布规律。

植物功能属性 是指把植物个体和生态系统结构、过程与功能联系起来的属性，反映植物对生态环境的响应和适应。

欢迎热爱生态学、关心全球环境变化的同学，加盟我们实验室，一起探索自然的奥秘！尤其鼓励有地学、植物（营养）学、化学和数学建模等学科背景或研究经历的同学，来实验室进行博士阶段的研究。

  奖励与荣誉

北京市“优秀本科毕业论文”指导教师（2019）；中国农业大学“优秀本科毕业论文”指导教师（2019/2014）；中国农业大学资环学院“优秀本科毕业论文”指导教师（2019/2012）；优秀班主任或优秀班主任标兵（2008/2009/2010/2011/2012），班级获评北京市优秀班集体（2008/2009）和北京先锋团支部（2008）。指导的博士生获北京市优秀博士毕业生（2019）。

  教学工作

主讲本科生课程《生态系统和全球变化》2019-

主讲本科生课程《土壤生态学》2013-2018

主讲研究生课程《全球生态学》2009-2012

参讲本科生课程《环境监测学》2011-2016

参讲研究生课程《环境科学技术》2007-2012

  主持项目

1.国家重点研发计划典型脆弱生态修复与保护专项“西部典型区域陆地生物多样性保护目标设定与评估示范”（2018.7-2021.7）

2.社会技术服务课题“清河流域水系生态环境调研”（2018.1-2019.12）

3.国家自然科学基金面上项目“中国北方森林植物多元素重吸收特征及其生态驱动机制”（2015-2018）

  学术成果

一、 发表文章：    

2019年：

1. Liu S, Yan Z, Chen Y, Zhang M, Chen J, Han W*. 2019. Foliar pH, an emerging plant functional trait: Biogeography and variability across northern China. Global Ecology and Biogeography 28:386-397.

2. Yan ZB, A Eziz, D Tian, XP Li, XH Hou, HY Peng, WX Han, YL Guo and JY Fang. 2019. Biomass allocation in response to nitrogen and phosphorus availability: insight from experimental manipulations of Arabidopsis thaliana. Frontiers in Plant Science. 10:598. doi:10.3389/fpls.2019.00598

3. Zhang MX, Luo Y, Yan ZB, Chen J, Eziz A, Li KH, Han WX*. 2019. Resorptions of ten mineral elements in leaves of desert shrubs and their contrasting responses to aridity. Journal of Plant Ecology 12(2):358-366.

4. Di Tian, Zhengbing Yan, Suhui Ma, Yuehong Ding, Yongkai Luo, Yahan Chen, Enzai Du, Wenxuan Han, Emoke Dalma Kovacs, Haihua Shen, Huifeng Hu, Jens Kattge, Bernhard Schmid, Jingyun Fang. 2019. Family-level leaf nitrogen and phosphorus stoichiometry of global terrestrial plants. Science China ▪ Life Sciences 62(8): 1047-1057.

5. Peng HY, Yan ZB, Chen YH, Zhao XJ, Han WX*. 2019. Effects of body size and root:shoot ratio on foliar nutrient resorption efficiency in Amaranthus mangostanus. American Journal of Botany 106(3): 363-370.

6. Fengzhan Geng, Kaihui Li, Xuejun Liu, Yanming Gong, Ping Yue, Yonggang Li, Wenxuan Han. 2019. Long-term effects of N deposition on N2O emission in an alpine grassland of Central Asia. Catena 182:104100

7. Qiqi Tan, Jiazhu Li, Zixun Chen, Ying Ma, Guoan Wang*, Yufu Jia, Hongyan Yao, Wenxuan Han*.2019. Clarifying the influence of temperature on variances in plant metallic nutrients through minimizing the effect of precipitation. Science of the Total Environment 646: 347-356.

8. Haojie Su, Yao Wu, Wulai Xia, Lei Yang, Jianfeng Chen, Wenxuan Han, Jingyun Fang, Ping Xie. 2019. Stoichiometric mechanisms of regime shifts in freshwater ecosystem. Water Research 149:302-310.

2018年：

1. Yindong Tong, Zhi Qiao, Xuejun Wang, Xueyan Liu, Guanyi Chen, Wei Zhang, Xin Dong, Zhengbing Yan, Wenxuan Han, Rong Wang, Menzhu Wang, Yan Lin. 2018. Human activities altered water N:P ratios in the populated regions of China. Chemosphere 210: 1070-1081.

2. Tang Xuli, Xia Zhao, Yongfei Bai, Zhiyao Tang, Wantong Wang, Yongcun Zhao, Hongwei Wan, Zongqiang Xie, Xuezheng Shi,Bingfang Wu, Gengxu Wang, Junhua Yan, Keping Ma, Sheng Du, Shenggong Li, Shijie Han, Youxin Ma, Huifeng Hu, Nianpeng He, Yuanhe Yang, Han Wenxuan, Hongling He,Guirui Yu, Jingyun Fang, Guoyi Zhou*. 2018. Carbon pools in China’s terrestrial ecosystems: New estimates based on an intensive field survey. Proceedings of the National Academy of Sciences of the United States of America 115(16):4021-4026.

3. Tang Zhiyao, Xu Wenting, Zhou Guoyi, Yongfei Bai, Jiaxiang Li, Xiuli Tang, Dima Chen, Qing Liu, Wenhong Ma, Gaoming Xiong, Honglin He, Nianpeng He, Yanpei Guo, Qiang Guo, Jiangling Zhu, Han Wenxuan, Huifeng Hu, Jingyun Fang, and Zongqiang Xie*. 2018. Patterns of plant carbon, nitrogen and phosphorus concentrations in relation to productivity in China's terrestrial ecosystems.Proceedings of the National Academy of Sciences of the United States of America 115(16):4033-4038.

4. Zhengbing Yan, Xiuping Li, Di Tian, Wenxuan Han, Xinghui Hou, Haihua Shen, Yalong Guo, Jingyun Fang. 2018. Nutrient addition affects scaling relationship of leaf nitrogen to phosphorus in Arabidopsis thaliana. Functional Ecology 32(12): 2689-2698.

5. Tian Ma, Shanshan Zhu, Zhiheng Wang, Dima Chen, Guohua Dai, Bowei Feng, Xiangyan Su, Huifeng Hu, Kaihui Li, Wenxuan Han, Chao Liang, Yongfei Bai & Xiaojuan Feng. 2018. Divergent accumulation of microbial necromass and plant lignin components in grassland soils. Nature Communications 9:3480.

6. Zhengbing Yan, Xinghui Hou, Wenxuan Han, Suhui Ma, Haihua Shen, Yalong Guo, Jingyun Fang. 2019. Effects of nitrogen and phosphorus supplies on stoichiometry of six elements in leaves of Arabidopsis thaliana. Annals of Botany 123(3):441–450. February 2019

7. Tan QQ, Li JZ, Chen ZX, Wang GA*, Jia YF, Yao HY, Han WX*.2018. Minimizing the effect of precipitation in clarifying the responses of leaf N and P stoichiometry to temperature. Environmental Pollution 243:404-409.

8. Tian D, Reich PB, Chen HYH, Xiang Y, Luo Y, Shen Y, Meng C, Han W, Niu S. 2018. Global changes alter plant multi-element stoichiometric coupling. New Phytologist 221(2): 807-817.

2017年：

1. Ethan Butler, Abhirup Datta, Habacuc Flores-Moreno, Ming Chen, Kirk R. Wythers, Farideh Fazayeli, Arindam Banerjee, Owen K. Atkin, Jens Kattge, Bernard Amiaud, Benjamin Blonder, Gerhard Boenisch, Ben Bond-Lamberty, Kerry Brown, Chaeho Byun, Giandiego Campetella, Bruno E Cerabolini, J. Hans C. Cornelissen, Joseph M. Craine, Dylan Craven, Franciska T. de Vries, Sandra Díaz, Tomas F. Domingues, Estelle Forey, Andres Gonzalez, Nicolas Gross, Wenxuan Han, Wesley N. Hattingh, Thomas Hickler, Steven Jansen, Koen Kramer, Nathan Kraft, Hiroko Kurokawa, Daniel C Laughlin, Patrick Meir, Vanessa Minden, Ulo Niinemets, Yusuke Onoda, Josep Peñuelas, Quentin Read, Fernando Valladares, Lawren Sack, Brandon Schamp, Nadejda A. Soudzilovskaia, Marko J. Spasojevic, Enio Sosinski, Peter E Thornton, Peter M. van Bodegom, Mathew Williams, Christian Wirth, Peter B. Reich. 2017. Mapping local and global variability in plant trait distributions. Proceedings of the National Academy of Sciences of the United States of America 114(51): E10937-E10946.  

2. Eziz A, Yan Z, Tian D, Han W, Tang Z, Fang J. 2017. Drought effect on plant biomass allocation: A meta analysis. Ecology & Evolution doi: 10.1002/ece3.3630.

3. Tian D, Yan Z, Niklas KJ, Han W, Kattge J, Reich PB, Luo Y, Chen Y, Tang Z, Hu H, Wright IJ, Schmid B, Fang J (2017). Global leaf nitrogen and phosphorus stoichiometry and their scaling exponent. National Science Review doi:10.1093/nsr/nwx142.

4. Zhang Meixia, Ji Chenjun, Zhu Jiangling, Wang Xiangping, Wang Donghui, Han Wenxuan*. 2017. Comparison of wood physical and mechanical traits between major gymnosperm and angiosperm tree species in China. Wood Science & Technology 51(6):1405-1419.

5. Yan Zhengbing, Tian Di, Han Wenxuan, Tang Zhiyao, Fang Jingyun. 2017. An assessment on the uncertainty of the nitrogen to phosphorus ratio as a threshold for nutrient limitation in plants. Annals of Botany 120(6):937-942.

6. 彭昀月 彭慧元 韩文轩. 2017. 豆科和非固氮植物氮磷利用效率研究. 中国农业大学学报 22(6):48-55.

7. Peng Yunyue, Peng Huiyuan, Han Wenxuan*. 2017. The comparison of nitrogen-and phosphorus-use efficiency between legume and non-nitrogen-fixing plant. Journal of China Agricultural University 22(6):48-55 (in Chinese)

2016年：

1. Zhengbing Yan, Wenxuan* Han, Pe?uelas Josep, Sardans Jordi, Elser James, Enzai Du, Reich Peter, Jingyun Fang. 2016. Phosphorus accumulates faster than nitrogen globally in freshwater ecosystems under anthropogenic impacts. Ecology Letters, 19: 1237-1246.

2. Huiyuan Peng, Yahan Chen, Yan Zhengbing Chen, Wenxuan* Han. 2016. Stage-dependent stoichiometric homeostasis and responses of nutrient resorption in Amaranthus mangostanus to nitrogen and phosphorus addition. Scientific Reports, 6: 37219.

3. Enzai Du, Vries WD, Wenxuan Han,Xuejun  Liu, Zhengbing Yan, Yuan Jiang. Imbalanced phosphorus and nitrogen deposition in China's forests. 2016. Atmospheric Chemistry and Physics, 16(13): 8571-8579.

4. Zhengbing Yan, Hanyue Guan, Wenxuan Han, Tingshen Han, Yalong Guo. Jingyun Fang. 2016. Reproductive organ and young tissues show constrained elemental composition in arabidopsis thaliana. Annals of Botany, 117(3): 431-439.

5. Yaoqi Li, Zhiheng Wang, Xiaoting Xu, Wenxuan Han, Qinggang Wang, Dongting Zou. 2016. Leaf margin analysis of Chinese woody plants and the constraints on its application to palaeoclimatic reconstruction: Leaf margin analysis of Chinese woody plants. Global Ecology & Biogeography, 25(12): 1401-1415.

6. Zhengbing Yan, Peng Li, Yahan Chen, Wenxuan Han, Jingyun Fang. 2016. Nutrient allocation strategies of woody plants: an approach from the scaling of nitrogen and phosphorus between twig stems and leaves. Scientific Reports, 6: 20099.

2015年：

1. Peng Li, Wenxuan Han, Chi Zhang, Di Tian, Xiaoniu Xu, Jingyun Fang. 2015. Nutrient resorption of Castanopsis eyrei varies at the defoliation peaks in spring and autumn in a subtropical forest, Anhui, China. Ecological Research, 30(1): 111-118.

2. Pin Li, Yuanhe Yang, Wenxuan Han, Jingyun Fang. 2014. Global patterns of soil microbial nitrogen and phosphorus stoichiometry in forest ecosystems. Global Ecology & Biogeography, 23: 979-987.

3. Zhengbing Yan, Namyoung Kim, Wenxuan Han, Yalong Guo, Tingshen Han, Enzai Du, Jingyun Fang. 2015. Effects of nitrogen and phosphorus supply on growth rate, leaf stoichiometry, and nutrient resorption of Arabidopsis thaliana. Plant and Soil, 388：147-155.

4. Yuanhe Yang, Pin Li, Honglin He, Xia  Zhao, Datta Arindam, Wenhong Ma, Ying Zhang, Xuejun Liu, Wenxuan Han, Wilson Maxwell, Jingyun Fang. 2015. Long-term changes in soil pH across major forest ecosystems in China: Soil pH dynamics in forests. Geophysical Research Letters, 42: 933-940.

5. Fanyun Yao, Yahan Chen, Zhengbing Yan, Peng Li, Wenxuan Han, Jingyun Fang. 2015. Biogeographic Patterns of Structural Traits and C:N:P Stoichiometry of Tree Twigs in China's Forests. Plos One, 10(2): e116391.

2014年：

1. Gong YM, Mohammat A, Liu XJ, Li KH, Christie P, Fang F, Song W, Chang YH, Han WX, Lu XT, Liu YY, Hu YK. 2014. Response of carbon dioxide emissions to sheep grazing and N application in an alpine grassland - Part 2: Effect of N application. Biogeosciences, 11: 1751-1757.

2. Liping Li, Stefan Zerbe, Wenxuan Han, Niels Thevs, Weiping Li, Ping He, Armin Schmitt, Yining Liu, Chengjun Ji. 2014. Nitrogen and phosphorus stoichiometry of common reed (Phragmites australis) and its relationship to nutrient availability in northern China. Aquatic Botany, 112: 84-90.

3. Chengjun Ji, Yuanhe Yang, Wenxuan Han, Yanfang He, Smith Jo, Smith Pete. 2014. Climatic and Edaphic Controls on Soil pH in Alpine Grasslands on the Tibetan Plateau: a Quantitative Analysis. Pedosphere, 24: 39-44.

4. Liping Li, Wenxuan Han, Thevs Niels, Xiuhong Jia, Chengjun Ji, Dongmei Jin, Ping He, Schmitt Armin, Cirella Giuseppe Tommaso, Zerbe Stefan. 2014. A Comparison of the Functional Traits of Common Reed (Phragmites australis) in Northern China: Aquatic vs. Terrestrial Ecotypes. Plos One, 9(2): e89063.

2013年：

1. Wenxuan Han, Luying Tang, Yahan Chen, Jingyun Fang. 2013. Relationship between the relative limitation and resorption efficiency of nitrogen vs phosphorus in woody plant. Plos One, 8: e83366.

2. Xuejun Liu, Ying Zhang, Wenxuan Han, Aohan Tang, Jianlin Shen, Zhenling Cui, Peter Vitousek, Jan Erisman, Keith Goulding, Peter Christie, Andreas Fangmeier, Fusuo Zhang. 2013. Enhanced nitrogen deposition over China. Nature, 494: 459-462.

3. Tang LY#, Han Wenxuan#, Chen YH, Fang JY. 2013. Resorption proficiency and efficiency of leaf nutrients in woody plants in eastern China. Journal of Plant Ecology, 6: 408-417.

4. Yahan Chen, Wenxuan Han, Luying Tang, Zhiyao Tang, Jingyun Fang. 2013. Leaf nitrogen and phosphorus concentrations of woody plants differ in responses to climate, soil and plant growth form. Ecography, 36: 178-184.

5. 严正兵, 金南瑛, 韩廷申, 方精云, 韩文轩*. 2013. 氮磷施肥对拟南芥叶片碳氮磷化学计量特征的影响. 植物生态学报, 37(6): 551-557.

2012年:

1. Wenxuan Han, Yahan Chen, FangJie Zhao, Luying Tang, Rongfeng Jiang, Fusuo Zhang. 2012. Floral, climatic and soil pH controls on leaf ash content in China’s terrestrial plants. Global Ecology & Biogeography, 21: 376-382.

2. Weiqi Shi, Guoan Wang*, Wenxuan Han*. 2012. Altitudinal Variation in Leaf Nitrogen Concentration on the Eastern Slope of Mount Gongga on the Tibetan Plateau, China. Plos One, 7: e44628.

3. Yuanhe Yang, Chengjun Ji, Wenhong Ma, Shifeng Wang, Shaopeng Wang, Wenxuan Han, Mohammat Anwar, Robinson David, Smith Pete. 2012. Significant soil acidification across northern China's grasslands during 1980s-2000s. Global Change Biology, 18: 2292-2300.

4. 何彦芳, 汤璐瑛, 陈雅涵, 李利平, 韩文轩*. 2012. 北京及周边区域植物叶片6种生理属性研究. 中国农业大学学报, 17(3)：81-87.

2011年:

1. Wenxuan Han, Jingyun Fang, Peter B Reich, F Ian Woodward, Zhiheng Wang. 2011. Biogeography and variability of eleven mineral elements in plant leaves across gradients of climate, soil and plant functional type in China. Ecology Letters, 14: 788-796.

2. Yuanhe Yang, Yiqi Luo, Meng Lu, Sch?del Christina, Wenxuan Han. 2011. Terrestrial C:N stoichiometry in response to elevated CO2 and N addition. Plant and Soil, 343: 393-400.

2010年:

1. Guo JH, Liu XJ, Zhang Y, JL Shen, Wenxuan Han, WF Zhang, P Christie, KWT Goulding, PM Vitousek, FS Zhang. 2010. Significant acidification in major Chinese croplands. Science, 327: 1008-1010.

2. 沈健林, 黄明丽, 唐傲寒, 汪斯淼, 韩文轩*, 刘学军. 2010. 葎草叶总氮和δ15N与大气活性氮气体的关系. 中国农业大学学报, 15(1): 84-88.

2009年:

1. 黄明丽, 韩文轩, 张福锁, 刘学军. 2009. 影响生物质能源植物产量和燃烧质量的农田管理因素分析. 农业工程学报, 25(2): 183-187.

2. 韩文轩, 吴漪, 汤璐瑛, 陈雅涵, 李利平, 贺金生, 方精云. 2009. 北京及周边地区植物叶的碳氮磷元素计量特征. 北京大学学报(自然科学版), 45(5): 855-860.

2008年:

1. Dali Guo, Harbin Li, Robert Mitchell, Wenxuan Han, Joseph Hendricks, Timothy Faheyand, and Ronald Hendrick. 2008. Fine root heterogeneity by branch order: exploring the discrepancy in root turnover estimates between minirhizotron and carbon isotopic methods. New Phytologist, 177:443-456.

2. 韩文轩, 方精云. 2008. 幂指数异速生长机制模型. 植物生态学报, 32(4): 951-960.

3. 韩文轩, 方精云. 2008. 植物种群的自然稀疏规律—-3/2还是-4/3？ 北京大学学报（自然科学版）,44(4): 661-66

2005年:

1. Wenxuan Han, Jingyun Fang, Dali Guo and Yan Zhang. 2005. Leaf nitrogen and phosphorus stoichiometry across 753 terrestrial plant species in China. New Phytologist, 168: 377-385.